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In an era where aliens have invaded and conquered feudal Tokyo, a young samurai seeks work as he can.

High schooler Rio Ozawa, falls for professor Kouki Ozawa. Though he reciprocates feelings, he's dying of cancer, explaining his distance. Rio convinces him to get risky surgery, even if he forgets her, to prolong their time together.

The Yorozuya gang returns to protect the country's shogun when the Shinsengumi police force finds itself in a crisis.

Kuno Totono, a curly-haired college student, visits Hiroshima and becomes embroiled in a mystery surrounding the inheritance of the Kariatsumari family's assets, using his intelligence and loquaciousness to unravel the case.